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We use deep Hubble Space Telescope imaging to study the evolutionary state of of the Virgo Cluster ultradiffuse galaxy (UDG) VCC 615. Using the tip of the red giant branch method, we pinpoint the galaxy's location with Virgo to be on the far side of the cluster, near the Virgo virial radius. When combined with the galaxy's measured line of sight velocity, we find the galaxy to be on an outbound orbit, having likely passed near the cluster core within the past billion years. Given the galaxy's largely undisturbed morphology, we argue that the galaxy has experienced no recent and sudden transformation into a UDG due to the cluster potential, but rather is a long-lived UDG whose relatively wide orbit and large dynamical mass protect it from stripping and destruction by Virgo cluster tides. Our Hubble imaging resolves individual globular clusters within VCC 615 down to a limiting half-light radius of 1 pc, allowing for a clean determination of the size distribution and luminosity function of the galaxy's globular cluster population. The total mass of the galaxy derived from the size of its globular cluster population is comparable to our previous work estimate from the mass from the globular cluster kinematics. Finally, we also derive the structural properties of the galaxy's nucleus and find them similar to the properties of ultracompact galaxies (UCDs) in the Virgo core, suggesting a dynamical link between UCDs and nucleated UDGs in cluster environments.